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Further problems arise as reproductive states and survival in wild populations are estimated based on imperfect and potentially biased observation processes, which might lead to flawed conclusions.

We developed a novel statistical framework that encapsulates the breeding life cycle of females, and s for incomplete information on female pregnancy and lactation and imperfect and biased recapture rates. We Women want sex Cherryfield this framework to longitudinal data on two sympatric, closely related bat species Myotis daubentonii and M.

Go to: 1.

Across taxa, experimental manipulation of reproductive effort has, for example, revealed that costs of reproduction lead to reduced survival e.

For example, costs of reproduction can become visible only under harsh environmental conditions e.

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Moreover, individuals may not experience any costs of reproduction or the costs will vary with individual quality e. This is because many studies suffer from imperfect i. The two species, however, show differences in their reproductive phenology timing of pregnancy and lactationfeeding habits see Old Orchard Beach horny sluts and hibernating behaviour M.

We develop a statistically rigorous framework that realistically describes the reproductive life cycle of females, while simultaneously incorporating uncertainty in the assessment of reproductive state and potentially biased recapture. Because bats are income breeders, reproduction and especially lactation should largely depend on the current food intake rather than fat reserves.

Reproduction, however, may cause females to allocate their current food intake in producing and feeding their young rather than in storing fat for themselves, such that females that reproduced Lady seeking sex NC Shiloh 27974 more likely to enter winter with lower fat reserves and suffer from reduced survival.

Furthermore, we predict that lactation as more energetically demanding and nearer to the onset of hibernation than pregnancy is more costly to survival than pregnancy only i. However, because M. Bats mainly take up residence in boxes in June after the bird breeding Seeking short and blondebut sometimes roost in boxes not used by birds prior to June.

Between andboxes were checked for bat occupancy between March and October. Over 26, box checks resulted in 1, recorded bat roosts containing between 1 and 45 individuals in boxes.

When captured for the first time, females were marked with a unique ID tag 2. We analysed data on M.

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The s of Mature petite boobs aged as juveniles when captured for the first time and thus of known age were M.

At temperate latitudes, the bat mating season occurs from autumn until spring, with both sexes storing sperm through winter hibernation Altringham, Myotis daubentonii specialize in feeding over calm open water, while M.

Breeders can be either successful lactating during the maternity period, SB or fail FB. When a female that has bred before i.

Green arrows represent possible state transitions between consecutive breeding seasons. Yellow lines within the shaded areas represent transitions that occur within breeding seasons.

However, depending on the timing and frequency of captures during the maternity period, this information relates with a different level of certainty to the true reproductive state of a female.

For example, if a female is encountered while pregnant, but not at a later stage, she could be either a successful or a failed breeder. The capture history of each female consists of a series of event codes, one code per year — Table 1 Eight possible events 0—7 that Housewives wants hot sex Chenango Forks used to construct a female encounter history.

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Each event relates to one or more possible underlying state s Early observation.